| List of Figures | xiii |
| Preface | xv |
| Acknowledgments | xvii |
| Introduction | xix |
| Purpose and Scope | xix |
| Physiographic Regions of South Carolina | xxi |
| Nature of the Flora | xxiii |
| What are Wildflowers? | xxiv |
| Conservation of Native Wildflowers | xxv |
| Collecting Native Wildflowers | xxvii |
| How to Use this Field Guide | xxviii |
| Origins of Plant Names | xxxi |
| Notes on Wildflower Photography | xxxiv |
| Rarity of Vascular Plants | xxxv |
| Pronunciation Guide to Scientific Names | xxxvii |
| Natural History of Selected Groups of Vascular Plants | 1 |
| Carnivorous Plants | 1 |
| Native Orchids | 5 |
| Poisonous Plants | 7 |
| Woody Vines | 8 |
| Epiphytic Vascular Plants | 10 |
| Saprophytic Vascular Plants and Mycotrophy | 11 |
| Parasitic Vascular Plants | 12 |
| Medicinal Plants and Folk Remedies | 13 |
| Edible Wild Plants | 17 |
| Naturalized Plants | 19 |
| Duckweeds | 21 |
| History of Field Botany in South Carolina | 23 |
| The Colonial Period: 1670-1785 | 24 |
| The Antebellum Period: 1785-1860 | 26 |
| The Civil War and the Postbellum Period: 1860-1900 | 31 |
| 1900-1952: the Beginning of Modern Field Botany | 33 |
| 1952-1983: the Wade Thomas Batson Era in South Carolina | 36 |
| Field Botany Today in South Carolina | 38 |
| Selected Topics on Natural History and Ecology | 39 |
| Succession in Natural Communities | 39 |
| Fire in the South Carolina Landscape | 41 |
| Carolina Bays of the Coastal Plain | 43 |
| Reproductive Strategies of Vascular Plants | 46 |
| Economic and Cultural Notes | 50 |
| Longleaf Pine and the Naval Stores Industry | 54 |
| Agriculture and Forestry: Effects on South Carolina's Physical Landscape | 56 |
| Marshes, Swamps, Peatlands, Bogs, and Fens | 65 |
| South Carolina's Natural Wildflower Communities | 69 |
| The Mountains | 69 |
| The Granitic Dome Community | 69 |
| Granitic Domes | 69 |
| The Spray Cliffs Community | 70 |
| Spray Cliffs | 70 |
| The Seepage Communities | 71 |
| Cataract Bogs | 71 |
| Montane Bogs | 71 |
| The Rocky Streamsides Community | 72 |
| Rocky Streamsides | 72 |
| The Deciduous Forest Communities | 73 |
| Cove Forests | 73 |
| Chestnut Oak Forests | 74 |
| Woodland Margins | 75 |
| Montane Oak-Hickory Forests | 76 |
| The Evergreen Forest Communities | 76 |
| Canada Hemlock Forests | 76 |
| Pine-Oak Heaths | 77 |
| The Piedmont | 78 |
| The Granitic Flatrocks Community | 78 |
| Granitic Flatrocks | 78 |
| The Rocky Shoals Community | 79 |
| Rocky Shoals | 79 |
| The Deciduous Forest Communities | 80 |
| Basic-Mesic Forests | 80 |
| Beech Forests | 81 |
| Oak-Hickory Forests | 82 |
| Piedmont Xeric Hardpan Forests | 83 |
| The Piedmont Springhead Seepage Forest Community | 84 |
| Piedmont Springhead Seepage Forests | 84 |
| The Bottomland Forest Communities | 85 |
| Bottomland Forests | 85 |
| The Fall-Line Sandhills | 86 |
| The Xeric Communities | 86 |
| Longleaf Pine-Scrub Oak Sandhills | 86 |
| Longleaf Pine-Turkey Oak Sandhills | 87 |
| The Sandhills Seepage Communities | 88 |
| Streamhead Pocosins | 88 |
| Herbaceous Seepages | 88 |
| Atlantic White-Cedar Forests | 89 |
| The Coastal Plain | 89 |
| The Pine Communities | 89 |
| Longleaf Pine Flatwoods | 89 |
| Pine/Saw Palmetto Flatwoods | 90 |
| Longleaf Pine Savannas | 91 |
| Longleaf Pine-Turkey Oak Xeric Ridges | 92 |
| The Sandy, Dry, Open Woodlands Community | 93 |
| Sandy, Dry, Open Woodlands | 93 |
| The Calcareous Forest Communities | 93 |
| Calcareous Bluff Forests | 93 |
| Wet, Flat, Calcareous Forests | 94 |
| The Bottomland Forest Communities | 94 |
| Bald Cypress-Tupelo Gum Swamp Forests | 95 |
| Hardwood Bottom Forests | 95 |
| Levee Forests | 96 |
| The Peatland Community | 97 |
| Pocosins | 97 |
| The Pond Cypress Communities | 98 |
| Pond Cypress Savannas | 98 |
| Pond Cypress-Swamp Gum Upland Swamps | 99 |
| The Freshwater Marsh Communities | 99 |
| Tidal Freshwater Marshes | 99 |
| Inland Freshwater Marshes | 100 |
| The Freshwater Floating Aquatics Community | 100 |
| Freshwater Floating Aquatics | 100 |
| The Depression Meadow Community | 101 |
| Depression Meadows | 101 |
| The Maritime Strand | 102 |
| The Maritime Communities | 102 |
| Coastal Beaches | 102 |
| Coastal Dunes | 102 |
| Maritime Forests | 103 |
| Salt Marshes | 104 |
| Salt Flats | 104 |
| Maritime Shell Forests | 105 |
| Species Descriptions and Color Plates | 107 |
| The Mountains | 107 |
| The Piedmont | 170 |
| The Fall-Line Sandhills | 225 |
| The Coastal Plain | 245 |
| The Maritime Strand | 343 |
| The Ruderal Communities | 365 |
| Field Guide to Natural Plant Communities | 409 |
| The Mountains | 409 |
| Greenville County | 409 |
| Caesar's Head State Park (Raven Cliff Falls Trail to Raven Cliff Falls Overlook) | 409 |
| Eva Russell Chandler Heritage Preserve | 409 |
| Watson Heritage Preserve (Mathews Creek Bog) | 410 |
| Mountain Bridge Trails | 411 |
| Table Rock State Park | 414 |
| Oconee County | 416 |
| Buzzard Roost Heritage Preserve | 416 |
| Sumter National Forest: Andrew Pickens District | 417 |
| Pickens County | 420 |
| Foothills Trail (Laurel Valley-Laurel Fork Falls) | 420 |
| Jocassee Gorges (Lower Eastatoe Creek Bluffs) | 422 |
| The Piedmont | 422 |
| Abbeville County | 422 |
| Sumter National Forest: Long Cane-Edgefield Districts | 422 |
| Aiken County | 424 |
| Savannah River Bluffs Heritage Preserve | 424 |
| Chester County | 425 |
| Landsford Canal State Park | 425 |
| Greenville County | 426 |
| Bunched Arrowhead Heritage Preserve | 426 |
| Lancaster County | 427 |
| Flat Creek Heritage Preserve (Forty Acre Rock) | 427 |
| Laurens County | 428 |
| Sumter National Forest: Tyger and Enoree Districts | 428 |
| McCormick County | 430 |
| Stevens Creek Heritage Preserve | 430 |
| Sumter National Forest: Long Cane-Edgefield Districts | 431 |
| Pickens County | 432 |
| Glassy Mountain Heritage Preserve | 432 |
| Union County | 434 |
| Sumter National Forest: Tyger and Enoree Districts | 434 |
| York County | 435 |
| Rock Hill Blackjacks Heritage Preserve | 435 |
| The Fall-Line Sandhills | 436 |
| Chesterfield County | 436 |
| Cheraw State Park (Hudsonia Flats) | 436 |
| Sandhills State Forest (Sugarloaf and Horseshoe Mountains) | 436 |
| Carolina Sandhills National Wildlife Refuge | 437 |
| Lexington County | 439 |
| Peachtree Rock Nature Preserve | 439 |
| Shealy's Pond Heritage Preserve | 439 |
| The Coastal Plain | 440 |
| Aiken County | 440 |
| Aiken Gopher Tortoise Heritage Preserve | 440 |
| Bamberg County | 441 |
| Cathedral Bay Heritage Preserve | 441 |
| Barnwell County | 442 |
| Craig Pond | 442 |
| Beaufort County | 442 |
| Victoria Bluff Heritage Preserve | 442 |
| Berkeley County | 443 |
| Francis Marion National Forest | 443 |
| Old Santee Canal Park | 444 |
| Wadboo Creek Calcareous Bluff Forest | 445 |
| Charleston County | 445 |
| Francis Marion National Forest | 445 |
| Santee Coastal Reserve | 446 |
| Clarendon County | 447 |
| Bennett's Bay | 447 |
| Dillion County | 447 |
| Little Pee Dee State Park | 447 |
| Dorchester County | 448 |
| Francis Beidler Forest | 448 |
| Georgetown County | 448 |
| Samworth Wildlife Management Area and Great Pee Dee River | 448 |
| Hampton County | 449 |
| Webb Wildlife Center | 449 |
| Horry County | 449 |
| Cartwheel Bay Heritage Preserve | 449 |
| Lewis Ocean Bay Heritage Preserve | 450 |
| Little Pee Dee River Heritage Preserve (Vaughan Tract) | 450 |
| Jasper County | 451 |
| Tillman Sand Ridge Heritage Preserve | 451 |
| Kershaw County | 451 |
| Savage Bay Heritage Preserve | 451 |
| Lee County | 452 |
| Longleaf Pine Heritage Preserve | 452 |
| Orangeburg County | 452 |
| Santee State Park | 452 |
| Sumter County | 453 |
| Woods Bay State Park | 453 |
| The Maritime Strand | 453 |
| Charleston County | 453 |
| Accreted Beach at Sullivan's Island | 453 |
| Beachwalker County Park | 454 |
| Colleton County | 454 |
| Edisto Beach State Park | 454 |
| St. Helena Sound Heritage Preserve (Otter Island) | 454 |
| Glossary | 457 |
Appendix 1 | South Carolina Herbariums | 471 |
Appendix 2 | Keys to Heartleafs, Rhododendrons, and Trilliums of South Carolina | 473 |
Appendix 3 | Illustrations of Plant Structures | 477 |
| General References | 481 |
| Literature Cited | 491 |
| Index of Scientific Plant Names | 495 |
| Index of Common Plant Names | 509 |
| General Index | 531 |